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Michelangelo's Sistine chapel frescoes are threatened by the effects of too many visitors, experts have warned on Wednesday, as the masterly painted ceiling celebrated its 500th anniversary.

"The anthropic pressure with dust, the humidity of bodies, carbon dioxide produced by perspiration can cause discomfort for the visitors and, in the long run, damage to the paintings," Antonio Paolucci, director of the Vatican Museums, wrote in the Vatican newspaper L'Osservatore Romano.